Thinning Practices in Southern Pines - With Pest Management Recommendations
United States Department of Agriculture, Forest Service, Technical Bulletin 1703, December 1985.
Growth of Trees and Stands
The principles of forest management are not greatly different from those affecting other agricultural crops. Trees, like other crops, require light, water, nutrients, space, and protection from insects and diseases. The fundamental growth processes are quite similar. The major difference is the length of time required to reach maturity. Given this difference, the economics of intensive management of a system as extensive as a forest has not always seemed favorable.
The growth potential of a tree is determined genetically, but actual growth is determined largely by the environment. Numerous environmental factors affect growth; of these, water, nutrients, and light intensity are most easily manipulated.
Height growth in the four major pines of the Southeast is indeterminate. Additional flushes (multinodal growth), particularly during midseason, reflect current soil moisture conditions. However, the initial terminal bud is formed in the year before extension, and the height growth from the extension of that bud is closely related to availability of soil water during the late summer of bud formation. If height increment is plotted against age, growth begins slowly at first, climbs more steeply, then flattens out (Prodan 1968). It is in these steeply climbing intermediate years, the grand period of growth between ages 10 and 30, that foresters try to regulate growth through thinning.
Diameter growth is also closely related to availability of soil water. Cambial cells begin dividing in early spring when soil water is not limited and stop in late summer when conditions are reversed. Across this ring of annual growth, early wood cells are abruptly followed by late wood cells whose greater density increases the specific gravity. Although the transition from early to late wood is not a well-understood process, , a decrease in soil water availability usually precedes the formation of late wood cells and a continued moisture deficit stops cell division. However, cell division may begin again in midsummer to late summer with increases in soil moisture, as evidenced by false annual rings. In addition, late wood cells continue to form until late summer or early fall if soil water is available (Moehring and Ralston 1967). It is partly through these biological principles that growth increments of individual trees can be regulated through thinning practices or stand density control.
Growth of stands is influenced by site quality, age, species, stocking level, and forestry practices.
A tree's environmental standing can be expressed through the concept of site quality. The site index is an integration of several environmental factors, but emphasizes the quality and quantity of soil nutrients and water. The rate of stand development increases with increased site index. Thus, the carrying capacity of a given unit of land for tree production increases with increasing site quality.
In an even-aged pure stand, the stages of development are similar throughout the stand at a given age, although more advanced stages are reached earlier on better quality sites than on low-quality sites. For similar stockings, a stand on a high-quality site will require thinning earlier than one on a low-quality site.
What are the implications of stocking and stand development? Proper stocking is a term commonly used but as difficult to apply as to define. First of all, proper stocking is the number of trees per acre that fully utilizes the site's potential to grow trees. It follows that a high-quality site has a higher carrying capacity and, if properly stocked, would carry more trees per acre than a low-quality site. Second, a given site may be properly stocked, once the carrying capacity is reached, with an initial spacing as low as 450 trees/acre or as high as 1,000 or more trees/acre. Third, the rate of diameter growth on individual stems and of stand development differs considerably with spacing. Both the rate of diameter growth and the age at which carrying capacity, in basal area, is reached are greater at wide spacings than at narrow spacings. It is these concepts of stocking, carrying capacity, and stand dynamics that form the biological basis for spacing and thinning to achieve management objectives.
The preceding discussion establishes the following premises:
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